In the version of this article initially published, there were errors in Figure 1. In Fig. 1f, due to a production error, the large panels for Ap4e1−/− mutant and control were interchanged and have been restored. In Fig. 1d, due to a submission error, large and small panels for Leprotl1−/− mutant and control were incorrect, and the smaller Fig. 1e Alpk3−/− control panel was also provided in error. The images have been replaced in the HTML and PDF versions of the article, and original and revised versions of Fig. 1d–f are shown below. (Figure presented.) (Figure presented.) Original Fig. 1d–f. Corrected Fig. 1d–f.